The only surviving witness of rapper Tupac Shakur's 1996 murder has been indicted, Fox News Digital can confirm.
Duane "Keffe D" Davis was indicted on the charge of open murder with use of a deadly weapon along with a gang enhancement, prosecutors revealed during a press conference Friday. In 2018, Davis made media statements that "reinvigorated" the investigation, police explained.
The charges were revealed Friday, hours after Davis, 60, was arrested while on a walk near his home, according to prosecutors.
Chief Deputy District Attorney Marc DiGiacomo said a grand jury had been seated in the case for "several months." The district attorney described Davis as the "on-ground, on-site commander" who "ordered the death" of Shakur.
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Police also revealed Davis obtained the gun used to kill Shakur from a "close associate." Davis was described as the "shot-caller" of a group of three suspects – all now deceased – and accused of orchestrating the plan to murder the rapper.
Davis has long been known to investigators and has himself admitted in interviews and in his 2019 tell-all memoir, "Compton Street Legend," that he was in the Cadillac when the gunfire erupted during the September 1996 drive-by shooting.
WATCH: POLICE RAID HOME IN CONNECTION WITH TUPAC SHAKUR'S MURDER
Police raided the suspect's wife's Nevada home on July 17. "Come out with your hands up and your hands empty!" law enforcement officers were heard yelling in the video as blue and red lights lit up the otherwise quiet neighborhood in the city of Henderson, which is about 20 miles from the Las Vegas Strip.
Authorities seized a Pokeball USB drive, an iPhone, three iPads (one with a cracked screen), four laptops, a tablet, a desktop computer, several external hard drives, copies of the book "Compton Street Legends," a Vibe magazine about Shakur and two "black tubs" of photos, according to the search warrant.
Police also searched for "notes, writings, ledgers and other handwritten or typed documents" about anything mentioning the murder of Shakur, the documents stated.
During Friday's press conference, authorities confirmed that they seized evidence using the search warrant that "corroborated" information they had collected during the investiagtion.
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Davis' nephew, Orlando Anderson, was considered the prime suspect in Shakur's 1996 murder. Anderson denied involvement before he was killed in a separate shooting in Compton, California, in 1998.
Shakur was fatally shot in a drive-by shooting on Sept. 7, 1996. The rapper was riding shotgun in Death Row Records co-founder Marion "Suge" Knight's black BMW when a white Cadillac pulled up alongside them while they were stopped at a light. In an interview done by Davis in 2018, he claimed everyone in the Cadillac that night was a member of the South Side Compton Crips gang.
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The gang was looking for Shakur, who had allegedly brawled with a member in the hours leading up to the fatal shooting.
Davis, who made the admission during the "Unsolved: The Murders of Tupac and the Notorious B.I.G." documentary, only revealed the gunshots came from the back seat.
Shakur was just 25 at the time. His fourth solo record, "All Eyez on Me," was still at the top of the charts with about 5 million copies sold.
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Fox News Digital's Chris Eberhart and The Associated Press contributed to this report.