BERLIN (AP) — Russian opposition leader Alexei Navalny, who is recovering in Germany after being poisoned in Russia by a nerve agent, accused Russian President Vladimir Putin of being behind the attack in comments released Thursday.

Navalny's supporters have frequently maintained that such an attack could have only been ordered at the top levels, although the Kremlin has repeatedly dismissed the accusations.

Navalny, a politician and corruption investigator who is Putin’s fiercest critic, was flown to Germany two days after falling ill on Aug. 20 on a domestic flight in Russia. He spent 32 days in the hospital, 24 of them in intensive care, before doctors deemed his condition had improved sufficiently for him to be discharged.

He has posted frequent comments online as his recovery has progressed, but in his first interview since the poisoning, he told Germany's Der Spiegel magazine that in his mind, “Putin was behind the attack,” in a German translation of his comments.

PUTIN TOUTS RUSSIAN CORONAVIRUS VACCINE, OFFERS IT TO UN STAFF FOR FREE

“I don't have any other versions of how the crime was committed," he said in a brief excerpt of the interview conducted in Berlin on Wednesday and to be released in full online later Thursday.

The Kremlin on Thursday said that “such accusations against the Russian president are absolutely groundless and unacceptable.”

“Some of these statements in the mentioned publication we consider offensive,” Kremlin spokesman Dmitry Peskov told reporters.

Peskov charged that there was information that “specialists” from the CIA were working with Navalny “these days” and giving him instructions.

“There is information that these instructors are working with him these days,” Peskov said. “Instructions the patient is receiving are obvious. We have seen such lines of behavior more than once.”

In this photo published by Russian opposition leader Alexei Navalny on his Instagram account on Monday, Sept. 21, 2020, Russian opposition leader Alexei Navalny and his wife Yulia pose for a photo in a hospital in Berlin. (Navalny Instagram via AP)

Earlier Thursday, Vyacheslav Volodin, speaker of the State Duma, Russia's lower parliament house, called Navalny “shameless” and “dishonorable" and accused the politician of working with “security services of Western countries.”

“Putin saved his life,” Volodin said in a statement released by the Duma. “Everyone, from pilots and doctors to the president, were genuinely saving him. Only a dishonorable person can make statements like this."

Navalny spent two days in a coma in a hospital in the Siberian city of Omsk, where Russian doctors said they found no trace of any poisoning, before being transported to Berlin for treatment. German chemical weapons experts determined that he was poisoned with the Soviet-era nerve agent Novichok — findings corroborated by labs in France and Sweden.

The nerve agent used in the attack was the same class of poison that Britain said was used on former Russian spy Sergei Skripal and his daughter in Salisbury, England, in 2018.

CLICK HERE TO GET THE FOX NEWS APP

German Chancellor Angela Merkel called the poisoning an attempted murder and she and other world leaders have demanded that Russia fully investigate the case.

Russia has bristled at the demands for an investigation, saying that Germany needs to share medical data in the case or compare notes with Russian doctors. Germany has noted that Russian doctors have their own samples from Navalny since he was in their care for 48 hours.

The Hague-based Organization for the Prohibition of Chemical Weapons has collected independent samples from Navalny for testing, but results haven't yet been announced.

German doctors have said the 44-year-old Navalny could make a full recovery, though haven't ruled out the possibility of long-term damage from the nerve agent.

Spiegel said Navalny was joking and alert in the interview, although his hands shook so much it was difficult for him to drink from a bottle of water. He also reiterated what his team has previously said – that he planned on returning to Russia when he was able to do so.

“My job now is to remain the guy who isn't scared,” he was quoted as saying. “And I'm not scared.”